They can. My vet told me to give some to Bilbo to help with his impaction. It makes the intestinal tract sticky and poo forms better. Only sometimes tho as its high in sugar!
They can, but I would be very careful about the sugar and the fact that it is quite high in acidity, which could cause mouth sores. I would only feed it in very small quantities. You may find that many piggies won't like it.
Sorry, i forgot to add that dangers of apples only effect some guinea pigs with a weakness in their lips. If your's have been eating apples for a long time and have never had any problems then it is fine to eat them.
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